1982 - DPhil. University of Oxford
1986-1991 - Royal Society University Research Fellow
1991-1993 - Lecturer, University of Manchester
1993-2001 - Lecturer, University of Cambridge
2001-2005 - Reader, University of Cambridge
2005 - Professor, University of Cambridge
Currently holds a joint appointment between the University of Cambridge and Cancer Research UKJoined Cancer Research UK Cambridge Research Institute in summer 2006.