1994 - MD PhD, Johns Hopkins University (Immunology), Baltimore, USA
Medical residency at Brigham and Women's Hospital
Medical oncology fellowship at the combined Dana-Farber/Partners Cancer Care programme
Postdoctoral fellowship with Tyler Jacks (MIT)
2002-2006 - Assistant Professor of Medicine, University of Pennsylvannia, USA.Joined the Cancer Research UK Cambridge Research Institute in summer 2006.