2004
Lecturer, University of Manchester School of Biological Science and CR-UK Career Development Fellow2001 - 2003
Post-doctoral research associate in the laboratory of Hans Clever, Academic Hospital Utrecht and Hubrecht Laboratories, The Netherlands
1999-2000
Wellcome Trust International Prize Travelling Fellow. Post-doctoral research performed in the laboratory of Hans Clevers, Academic Hospital Utrecht, The Netherlands
1998
PhD University of Glasgow, Faculty of Medicine. Research performed at CR-UK Beatson laboratories under the supervision of Dr Donald M Black.
1994
BA (Hons) University of Cambridge, Natural Sciences.